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WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
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Cyperus esculentus L.

Accepted
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
Cyperus esculentus L.
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Cyperus esculentus L.
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🗒 Synonyms
synonymChlorocyperus aureus (K.Richt.) Palla ex Kneuck.
synonymChlorocyperus phymatodes (Muhl.) Palla
synonymCyperus aureus subsp. esculentus (L.) Nyman, nom. superfl.
synonymCyperus aureus Ten. [Illegitimate]
synonymCyperus aureus Ten., nom. illeg.
synonymCyperus bahiensis Steud.
synonymCyperus buchananii Boeckeler
synonymCyperus bulamensis Steud.
synonymCyperus callistus Ridl.
synonymCyperus chrysostachys Boeckeler
synonymCyperus cubensis Steud.
synonymCyperus damiettensis A.Dietr.
synonymCyperus esculentus f. angustispicatus (Britton) Fernald
synonymCyperus esculentus f. evolutus C.B.Clarke
synonymCyperus esculentus f. macrostachyus (Boeckeler) Fernald
synonymCyperus esculentus f. princeps C.B.Clarke
synonymCyperus esculentus subsp. aureus K.Richt.
synonymCyperus esculentus var. angustispicatus Britton
synonymCyperus esculentus var. cyclolepis Boeckeler ex Kük.
synonymCyperus esculentus var. esculentus
synonymCyperus esculentus var. heermannii (Buckley) Britton
synonymCyperus esculentus var. helodes (Schrad. ex Nees) C.B.Clarke
synonymCyperus esculentus var. leptostachyus Boeckeler
synonymCyperus esculentus var. lutescens (Torr. & Hook.) Kük. ex Osten
synonymCyperus esculentus var. macrostachyus Boeckeler
synonymCyperus esculentus var. phymatodes (Muhl.) Kük.
synonymCyperus esculentus var. sativus Boeckeler
synonymCyperus esculentus var. sprucei C.B.Clarke
synonymCyperus fulvescens Liebm.
synonymCyperus gracilescens Schult.
synonymCyperus gracilis Link [Illegitimate]
synonymCyperus heermannii Buckley
synonymCyperus helodes Schrad. ex Nees
synonymCyperus hydra Kunth [Illegitimate]
synonymCyperus hydra Kunth, nom. illeg.
synonymCyperus lutescens Torr. & Hook.
synonymCyperus melanorhizus Delile
synonymCyperus nervosus Bertol.
synonymCyperus officinalis T.Nees
synonymCyperus pallidus Savi [Illegitimate]
synonymCyperus pallidus Savi, nom. illeg.
synonymCyperus phymatodes Muhl.
synonymCyperus phymatodes var. heermannii (Buckley) S.Watson
synonymCyperus repens Elliott
synonymCyperus ruficomus Buckley
synonymCyperus sieberianus Link
synonymCyperus tenoreanus Schult.
synonymCyperus tenoreanus Schult. & Schult.f.
synonymCyperus tenorei C.Presl
synonymCyperus tenorianus Roem. & Schult.
synonymCyperus tuberosus Pursh [Illegitimate]
synonymCyperus tuberosus Pursh, nom. illeg.
synonymCyperus variabilis Salzm. ex Steud.
synonymPterocyperus esculentus (L.) Opiz
synonymPycreus esculentus (L.) Hayek
🗒 Common Names
Afrikaans
  • Geeluintjie
Comorian
  • Makunu
Créole Maurice
  • Souchet comestible
Créole Réunion
  • Gros zoumine
Créole Seychelles
  • Laiche
  • Herbe oignon
  • Lerb zonnyon
English
  • Yellow nutsedge, Tiger nut, Earth almond, Nutgrass, Tigernut, Watergrass, Yellow nut grass
French
  • Amande de terre, Noix tigrée (Antilles)
Malgache
  • Karepoka
  • Tsingetsetse
Other
  • N'drawe, N'drawe n'drume (Shimaore, Mayotte)
  • Androy, Androy lahy (Kibushi, Mayotte)
Spanish; Castilian
  • Cebollín, Chufa, Chufa commún, Coco, Coquillo, Coquillo amarillo, Coquito, Coyolillo, Juncia, Juncia avellanada
📚 Overview
Overview
Brief
Code

CYPES

Growth form

sedge

Biological cycle

vivacious

Habitat

terrestrial

Wiktrop
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Lovena Nowbut
StatusUNDER_CREATION
LicensesCC_BY
References
    Diagnostic Keys
    Description
    Global description
     
    Cyperus esculentus is a vivacious sedge that grows in small tuft with a single flowering axis, but connected to many progeny by the rhizomes. At the base of a mother plant is a spherical and black mother tuber. The inflorescence consists of an umbel whose floral axis have golden yellow spikelets at the top.
     
    First leaves
     
    The first leaves are linear, acute apex, 2 to 10 cm long and 4 mm wide. The section of blade form a wide V. At the base of the blade is a short trigonal sheath. The leaves are arranged in a tristichous fashion. The seedling can be connected to a mother plant via a rhizome or isolated. The seedling is then protruded from a spherical and black mother tuber, more or less deep that develops a vertical axis, scaly at the underground part, turning into small foliage tuft above ground.
     
    General habit
     
    Plant with tufted growth habit. This tuft is constituted of the interlocking of the leaf sheaths in a tristichious way. Each tuft develops only one erect floral axis. The plant can measure 30 to 80 cm high. A mother plant is connected to the progeny by superficial rhizomes.
     
    Underground system
     
    The roots are fibrous. They are simple, filiform and very numerous, formijng an important root hairs. The underground part of C. esculentus is characterized by the presence of a black spherical mother tuber at the base of the main axis but also by white horizontal rhizomes, with few scales and emerging a few centimeters away to give a new plant. The mother plant is also develop deeper rhizomes at the end of which there is a single, white, spherical tuber, slightly scaly. These tubers are up to 1 cm in diameter. They have a sweet smell and taste.
     
    Stem
     
    The aerial stem is the flowering axis. It is full with triangular section and sharp angles. It is 3 to 6 mm large. It is completely smooth and bright green.
     
    Leaf
     
    The leaves are linear, sometimes as long as or longer than the stem. They are located in the lower part of the stem and arranged in a tristichous way. The sheath is trigonal. The blade is 5 to 6 mm long. It has a wide V shape in section. The margin is scabrous at the end part of the leaf blade. The upper side is glossy light green while the underside is paler. Both sides are glabrous.
     
    Inflorescence
     
    The flowers are assembled as flat spikelets arranged on top of the floral axis of the umbel, which makes up the inflorescence. This umbel can be simple or compound. It is subtended by 3 to 5 leafy bracts,5 to 25 cm long. The umbel has many long and glabrous floral axis, 2 to 10 cm. Each held along the 2 to 3 cm terminals, 5 to 30 spikelets arranged perpendicularly to the axis.
     
    Flower
     
    The spikelets are linear, they are 1 to 3 cm long and 1 to 1.5 mm wide. They are flat because the flowers are arranged in a distichous alternate way along the rachis. A spikelet consists of 8 to 14 flowers. The glumes are 3 mm long. In the spikelet, the top of a glume reach the middle of the previous glume. The top is slightly notched and very shortly mucronate. The glume is marked by 3 or 4 strong longitudinal striations. It is golden yellow when ripe with a lighter papery margin. The ovary is surmounted by a long trifid style, exceeding the glumes.
     
    Fruit
     
    The fruit is an trigonal to obovoid achene, slightly flattened and with rounded corners. It is 1.5 mm long and 0.7 mm wide. It is greyish in color and topped by a trifid style. The seed coat is decorated with small alveoli.

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      No Data
      📚 Natural History
      Life Cycle

      Life cycle

      Vivacious
      Vivacious

      Northern Cameroon: The emergence of young plants of Cyperus esculentus is carried out throughout the humid period of soil. It is favored by weeding, which breaks the dormancy of the tubers by separating them from the mother plant, Flowering is induced by long days and takes place between July and October, 4 to 8 weeks after emergence. Some populations do not flower, especially the cultivated populations. The plants wither at the beginning of the dry season, depending on the drying of the soil. 

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        Reproduction
        Cyperus esculentus is a vivacious species which is mostly propagated vegetatively. On one side, the horizontal rhizomes are located under the ground surface, directly give rise to new plant. On the other hand, rhizomes heading deep, end with a more or less dormant tuber, from which grows a new plant after the dry season. A mother plant from an initial tuber can produce 36 saplings and 339-progeny tubers in 16 weeks of vegetation. C. esculentus is also propagated by seed.

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          Morphology

          Growth form

          Tuft plant with narrow leaves
          Tuft plant with narrow leaves

          Leaf type

          Grass or grass-like
          Grass or grass-like

          Latex

          Without latex
          Without latex

          Stem section

          Triangular
          Triangular

          Root type

          Tubers
          Tubers
          Fibrous roots
          Fibrous roots

          Stipule type

          No stipule
          No stipule

          Leaf attachment type

          Cyperaceae leaf
          Cyperaceae leaf

          Achene type

          Achene trigonous
          Achene trigonous

          Lamina base

          sheathing the triangular stems
          sheathing the triangular stems

          Lamina margin

          scabrous
          scabrous
          entire
          entire

          Lamina apex

          attenuate
          attenuate

          Simple leaf type

          Lamina linear
          Lamina linear

          Flower color

          Yellow
          Yellow
          Orange
          Orange

          Inflorescence type

          Digitate racemes
          Digitate racemes
          Umbel
          Umbel

          Life form

          Geophytic plant
          Geophytic plant
          Sedge leaf
          Sedge leaf
          Look Alikes
          Cyperus esculentus can be easily confused in the vegetative stage with C. rotundus. An adult C. esculentus emits rhizomes comprising of a single, terminal, spherical tuber, with a sweet taste, whereas C. rotundus produce ellipsoidal tubers with fibrous walls of dark color arranged in strings along branched rhizomes. In addition, C. rotundus tubers as well as the base of the plant emit a strong peppermint odor. The inflorescence of C. esculentus is formed from an umbel whose floral axis have golden yellow spikelets on the top, arranged perpendicular to the rachis.

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            Ecology

            Northern Cameroon: Cyperus esculentus is found in all soils with a pH between 5 and 7. This species is usually more abundant on sandy loam alluvial soils along river. It is very tolerant to high soil moisture. It is often associated with C. rotundus, but in the most humid stations, it tends to replace C. rotundus and Cynodon dactylon. It tolerates shade in crop cultivations.
            Comoros: ruderal plant that is found mostly in areas exposed to the sun, at low and medium altitude.
            Madagascar: Species quite common in dry crops, which settles in relatively rich and moist enough soils, down slope or in the alluvial plains.
            Mauritius: A weed of crops, it is found mainly in the wet and cool stations.
            Mayotte: C. esculentus is an uncommon weed, which occurs in fruit and vegetable crops in the rainy northern region of the island.
            Reunion: rare species but distributed over the entire circumference of the island and can reach up to 1000 m altitude
            Seychelles: This species is observed in heavily irrigated crops in wet fields and heavy soils. It tolerates very wet soils, but grows well on all soil types.

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              No Data
              📚 Habitat and Distribution
              Description

              Geographical distibution

              Madagascar
              Madagascar
              Reunion Island
              Reunion Island
              Comoros
              Comoros
              Mauritius
              Mauritius
              Seychelles
              Seychelles
              Origin

              Cyperus esculentus is native to both North America, and South America, as well as Eurasia and Africa.

              Worldwide distribution

              Widely distributed in India, Indochina, Madagascar, the Mediterranean and South Europe.

              dummy
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              StatusUNDER_CREATION
              LicensesCC_BY_SA
              References
                No Data
                📚 Occurrence
                No Data
                📚 Demography and Conservation
                Risk Statement

                Global harmfulness

                Tubers and roots being interconnected to each other to a depth of 50 cm (19.68 inch) or more. The tubers are connected by fragile roots that are extremely prone to snapping when pulled on, so it is extremely difficult to remove Cyperus esculentus with its entire root system intact. The plant will regenerate completely if even a single tuber is left in place. Tubers typically survive up to ~ 3-4 years.

                Local harmfulness

                Benin: frequent and generally abundant.
                Burkina Faso: frequent and scarce.
                Northern Cameroon: C. esculentus is a local major weed.
                Chad: frequent and generally abundant.
                Comoros
                : A weed present in banana plantations, cassava and sugar cane.
                Ivory Coast: common and scarce.
                Ghana: frequent and generally abundant.
                Kenya: Cyperus esculentus is frequent and usually abundant.
                Madagascar: A weed of low frequency, but sometimes locally abundant as small spots more or less widespread, especially in corn and bean crops.
                Mali: rare but abundant when present.
                Mauritius important weed of crops may be highly harmful.
                Mayotte: C. esculentus is an infrequent weed, present in 1% of cultivated plots, but can be locally abundant in fruit and vegetable crops.
                Nigeria: frequent and generally abundant.
                New Caledonia: The species is reported since 1850 and is the main weed in vegetable gardening. It has remarkable plasticity. However environmental factors affect its development, the humidity is therefore essential to its reproduction and growth. Under certain conditions, it can therefore also become an invasive species of degraded or planted forage crops. But it does not tolerate competition for light from other plants and tends to disappear when the vegetation grows. The tubers remain dormant in the soil until the next tillage.
                Uganda: common and generally abundant.
                Reunion: A weed infrequent, occasionally encountered in arboriculture, sugar cane and especially vegetable cultures because it is favored by rotovator tillage. It develops by multiplication of tubers, leading in certain situations collection, 30-50% of the plot.
                Senegal: rare but abundant when present.
                Seychelles: serious or principal weed of fruit crops, vegetable and tubers.
                Tanzania: frequent and generally abundant.
                West Indies: Cyperus esculentus is favoured by shallow cultivation operations, which separate and disperse the tubers. It is not sensitive to pre-emergence herbicides and not very sensitive to post-emergence contact or systemic herbicides. Because of its ability to multiply vegetatively throughout the growing season, this species becomes invasive very quickly.

                 

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                  No Data
                  📚 Uses and Management
                  Uses

                  Food: In Northern Cameroon, Cyperus esculentus is cultivated for its tubers, used in human food. The tuber flour of C. esculentus is a source of carbohydrates and some mineral elements such as iron and calcium.

                   

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                    Management
                    Global control
                    Chemical control: As Cyperus rotundus, C. esculentus is relatively sensitive to 2, 4-D or MCPA at high doses. The tops of young plants are destroyed and regrowth may be delayed but the tubers themselves are unaffected. In young sugarcane, this level of control may retard weed growth long enough so that culture can settle and make enough shade on later emergence. In most cultures, these effects are not durable enough. Among other chemicals foliar absorption, high dose glyphosate (preferably in a double application, first to 2000 g / ha and 1000 g / ha regrowth) is effective and can be used either before planting or as a directed spray. When its use was allowed, paraquat destroys the top of the plant, but it had to be applied several times to 4 weeks apart to achieve a lasting effect.
                     
                    Biological control: In traditional rice-growing areas in India C. rotundus is controlled using pigs. Usually, the pigs are fed with vegetable waste and they look for tubers in wastelands. The pigs love these delicious and sweet potatoes. They can easily extract the tubers even if the soil is hard. Usually the field is watered and then the pigs are introduced on the plot. An animal may recover 2-4 kg of tubers per day and sixty animals can clean the tubers from 1 ha in one day.
                     
                    Tips for weeding perennial cyperaceae irrigated rice and lowland in Africa, visit: http://portal.wikwio.org/document/show/29
                     
                    Local control
                     
                    Madagascar: Tillage or pulling (very laborious) separates the mother plant tubers, releasing progeny plant of the dominance of the mother plant, thus contributing to their rapid multiplication. Tolerant to most pre-emergence herbicides. Sensitive to 2, 4-D and glyphosate, but in difficult application conditions on crops. It is best to control these weeds (glyphosate at 2160 g / ha) at the end of growing season to prepare for the following year. However, cyperaceae do not tolerate shade. A good ground cover that delays (but does not stop) their emergence and a dense planting of a crop that quickly covers the ground allow to properly control them.
                    Wiktrop
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                      No Data
                      📚 Information Listing
                      References
                      1. Le Bourgeois, T., Carrara, A., Dodet, M., Dogley, W., Gaungoo, A., Grard, P., Ibrahim, Y., Jeuffrault, E., Lebreton, G., Poilecot, P., Prosperi, J., Randriamampianina, J.A., Andrianaivo, A.P., Théveny, F. 2008. Advent-OI : Principales adventices des îles du sud-ouest de l'Océan Indien.V.1.0. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom.
                      1. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                      1. Le Bourgeois, T. and H. Merlier (1995). Adventrop - Les adventices d'Afrique soudano-sahélienne. Montpellier, France, Cirad. 640 p.
                      2. Grossard, F., Le Bourgeois, T., Dumbardon-Martial, E. & Gervais, L. 2013. Adventilles - Guadeloupe & Martinique - Les adventices des Antilles françaises. Abymes, Guadeloupe, France, Les éditions du CTCS Guadeloupe. 195 p.
                      3. Fournet, J. 2002. Flore illustrée des phanérogames de Guadeloupe et de Martinique. Montpellier, France, Cirad, Gondwana éditions.
                      4. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:300668-2
                      1. Husson, O., H. Charpentier, F.-X. Chabaud, K. Naudin, Rakotondramanana et L. Séguy (2010). Flore des jachères et adventices des cultures. Annexe 1 : les principales plantes de jachères et adventices des cultures à Madagascar. In : Manuel pratique du semis direct à Madagascar. Annexe 1 - Antananarivo : GSDM/CIRAD, 2010 : 64 p.
                      1. http://www.issg.org/database/species/ecology.asp?si=1448&fr=1&sts=sss&lang=FR
                      1. http://idao.cirad.fr/SpecieSheet?sheet=oswald/species/c/cypes/cypes_fr.html
                      1. TAMES, R.S., GETSO, M.D.V. & VIEITEZ, E., 1973. Growth substances isolated from tubers of Cyperus esculentus var aureus. Physiology of Plants 28, 195-200;
                      2. G. W. Ivens (1989). Eastern Africa weeds control. Oxford University press, Nairobi. 17p;
                      3. Troupin G. (1987). Flore du Rwanda, spermatophytes (Volume IV), Musée Royal de l'Afrique centrale, Tervuren, Belgique. Musée Royal de l'Afrique centrale, Tervuren, Belgique.438p.
                      1. Huat, J., Nagy, M., Carpente, A., Schwartz, M., Le Bourgeois, T. & Marnotte, P. 2021. Guide de la flore spontannée des agrosystèmes de Mayotte. Montpellier, Cirad. 150 p.
                      2. Le Bourgeois, T. and H. Merlier (1995). Adventrop - Les adventices d'Afrique soudano-sahélienne. Montpellier, France, Cirad. 640 pp.
                      1. Blanfort V., Desmoulins F., Prosperi J., Le Bourgeois T., Guiglion R. & Grard P. 2010. AdvenPaC V.1.0 : Adventices et plantes à conflit d'intérêt des Pâturages de Nouvelle-Calédonie. IAC, Cirad, Montpellier, France, Cédérom. http://idao.cirad.fr/applications.
                      1. http://www.hear.org/pier/species/cyperus_esculentus.htm
                      Information Listing > References
                      1. Le Bourgeois, T., Carrara, A., Dodet, M., Dogley, W., Gaungoo, A., Grard, P., Ibrahim, Y., Jeuffrault, E., Lebreton, G., Poilecot, P., Prosperi, J., Randriamampianina, J.A., Andrianaivo, A.P., Théveny, F. 2008. Advent-OI : Principales adventices des îles du sud-ouest de l'Océan Indien.V.1.0. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom.
                      2. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                      3. Le Bourgeois, T. and H. Merlier (1995). Adventrop - Les adventices d'Afrique soudano-sahélienne. Montpellier, France, Cirad. 640 p.
                      4. Grossard, F., Le Bourgeois, T., Dumbardon-Martial, E. & Gervais, L. 2013. Adventilles - Guadeloupe & Martinique - Les adventices des Antilles françaises. Abymes, Guadeloupe, France, Les éditions du CTCS Guadeloupe. 195 p.
                      5. Fournet, J. 2002. Flore illustrée des phanérogames de Guadeloupe et de Martinique. Montpellier, France, Cirad, Gondwana éditions.
                      6. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:300668-2
                      7. Husson, O., H. Charpentier, F.-X. Chabaud, K. Naudin, Rakotondramanana et L. Séguy (2010). Flore des jachères et adventices des cultures. Annexe 1 : les principales plantes de jachères et adventices des cultures à Madagascar. In : Manuel pratique du semis direct à Madagascar. Annexe 1 - Antananarivo : GSDM/CIRAD, 2010 : 64 p.
                      8. http://www.issg.org/database/species/ecology.asp?si=1448&fr=1&sts=sss&lang=FR
                      9. http://idao.cirad.fr/SpecieSheet?sheet=oswald/species/c/cypes/cypes_fr.html
                      10. TAMES, R.S., GETSO, M.D.V. & VIEITEZ, E., 1973. Growth substances isolated from tubers of Cyperus esculentus var aureus. Physiology of Plants 28, 195-200;
                      11. G. W. Ivens (1989). Eastern Africa weeds control. Oxford University press, Nairobi. 17p;
                      12. Troupin G. (1987). Flore du Rwanda, spermatophytes (Volume IV), Musée Royal de l'Afrique centrale, Tervuren, Belgique. Musée Royal de l'Afrique centrale, Tervuren, Belgique.438p.
                      13. Huat, J., Nagy, M., Carpente, A., Schwartz, M., Le Bourgeois, T. & Marnotte, P. 2021. Guide de la flore spontannée des agrosystèmes de Mayotte. Montpellier, Cirad. 150 p.
                      14. Le Bourgeois, T. and H. Merlier (1995). Adventrop - Les adventices d'Afrique soudano-sahélienne. Montpellier, France, Cirad. 640 pp.
                      15. Blanfort V., Desmoulins F., Prosperi J., Le Bourgeois T., Guiglion R. & Grard P. 2010. AdvenPaC V.1.0 : Adventices et plantes à conflit d'intérêt des Pâturages de Nouvelle-Calédonie. IAC, Cirad, Montpellier, France, Cédérom. http://idao.cirad.fr/applications.
                      16. http://www.hear.org/pier/species/cyperus_esculentus.htm

                      Etude floristique et phytoécologique des adventices des complexes sucriers de Ferké 1 et 2, de Borotou-Koro et de Zuenoula, en Côte d'Ivoire

                      Thomas Le Bourgeois
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                        🐾 Taxonomy
                        📊 Temporal Distribution
                        📷 Related Observations
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                        WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areasWIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
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